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[00]  Tamaricaceae (Draft: R. Elven)

THIS DRAFT HAS NOT BEEN DISCUSSED
Myricaria Desv. (1825), Ann. Sci. Nat. 4: 348, with M. germanica (L.) Desv. (1825), Ann. Sci. Nat. 4: 349 [basionym: Tamarix germanica L. (1753), Sp. Pl. 271] nearly reaches the Arctic in N Norway (Porsanger and Lebesby). Could perhaps be included as a border case. (Elven)

68  Elatinaceae (Draft: R. Elven)

THIS DRAFT HAS NOT BEEN DISCUSSED

68.1  Elatine L. (1753) Sp. Pl. 367.
68.1.1  Elatine triandra Schkuhr (1791), Handb. 1: 345., coll.

S E. triandra Schkuhr var. gracilis ***

2n=

2nD


G CAN

Comments:

(1) Mapped from one site on the arctic coast of Yukon Territory by Cody (1996). (Elven)

69  Elaeagnaceae (Draft: R. Elven)

THIS DRAFT HAS NOT BEEN DISCUSSED
69.1  Elaeagnus L. (1753), Sp. Pl. 121.
69.1.1  Elaeagnus commutata Bernh. ex Rydb. (1918), Fl. Rocky Mts. 582.

S

T [Described from USA: "In the extensive plains on the banks of the Missouri".]



2n=

2nD


G CAN?

Comments:

(1) Approaches or reaches the arctic parts in S Hudson Bay area (Porsild & Cody 1980). Scoggan (1979): "20 mi SW of York Factory ... and northernmost Ont[ario]". An acceptable border case. (Elven)

69.2  Shepherdia Nutt. (1818), Gen. Amer. 2: 240.
69.2.1  Shepherdia canadensis (L.) Nutt. (1818), Gen. Amer. 2: 240.

B Hippophae canadensis L. (1753), Sp. Pl. 1024.

S

T [Described from "Canada".]



2n= 22 (2x).

2nD Cooper (1932); Löve & Löve in Löve (1975d).

G ALA CAN

Comments:



70  Onagraceae (Draft: A.K. Skvortsov)

THIS DRAFT HAS NOT BEEN DISCUSSED

Comments:

(1) We should attain some consistency in how broad or narrow species and genera are defined throughout the checklist. Skvortsov perhaps applies a slightly broader definition of species than many other draft authors. I have therefore tentatively raised a few of the taxa that Skvortsov includes only in synonymy. (Elven)

(2) Oenothera L. (1753), Sp. Pl. 346, is not mentioned in Skvortsov's draft but is reported from the Arctic: a species of the O. biennis L. (1753), Sp. Pl. 346, complex is mapped from the SW side of Hudson Bay (Hultén & Fries 1986 map 1354). Probably not stable and therefore not included formally. (Elven)

(3) Another genus   Circaea   is added below. (Elven)

70.1  Epilobium L. (1753), Sp. Pl. 347.
The Epilobium palustre aggregate (E. palustre, E. tundrarum)

Comments:

(1) Skvortsov in Tolmachev & Yurtsev (1980, Fl. Arct. URSS 8) and in draft synonymises E. tundrarum with E. palustre whereas Flora Europaea 2 (1968) and Löve & Löve (1975) keep it as a separate species. I have tentatively followed the latter authorities. Treatment as a subspecies may be more appropriate. (Elven)
70.1.1  Epilobium palustre L. (1753), Sp. Pl. 348.

S

2n= 36 (6x?).



2nD Löve & Löve (1975) list several counts, three as arctic.

G ICE NOR RUS SIB RFE ALA CAN GRL

Comments:

(1) Check whether E. leptophyllum Raf. (1808), Med. Repos. New York 2, 5: 353, reaches the Arctic SE of Hudson Bay. Not acc. to Porsild & Cody (1980). (Elven)


70.1.2  Epilobium tundrarum Sam. (1922), Bot. Not. 1922: 264.

S

2n=



2nD

G RUS


Comments:

(1) Reported by Samuelsson (1922) from Waigach and Polar Ural, probably as a very local race. (Elven)



WARNING! Should probably be reduced to a subspecies of E. palustre.
70.1.3  Epilobium davuricum Fisch. ex Hornem. (1819), Hort. Hafn., Suppl. 44.

S

Comments:



(1) Skvortsov synonymises E. arcticum with E. davuricum, with comment. It is here accepted as a subspecies as usually done in N Europe and N America. (Elven)
70.1.3.1  Epilobium davuricum Fisch. subsp. davuricum

S

2n= 36 (6x?).



2nD Knaben (1950 Norw); Engelskjøn & Knaben (1971 Norw); Zhukova & Tikhonova (1971).

G NOR RUS SIB ALA CAN GRL? (E)

Comments:

(1) The identity of E Greenland plants needs some verification. (Elven)


70.1.3.2  Epilobium davuricum Fisch. subsp. arcticum (Sam.) P.H. Raven (1968), Feddes Repert. 79: 61.

B E. arcticum Sam. (1922), Bot. Not. 1922: 260.

S

2n=


2nD

G RUS SIB RFE ALA CAN GRL

Comments:
70.1.4  Epilobium alsinifolium Vill. (1779), Prosp. Hist. Pl. Dauphiné 45.

S

2n= 36 (6x?).



2nD Löve & Löve (1975) list three counts, one as arctic.

G ICE NOR RUS GRL

Comments:

(1) Added to Skvortsov's draft because it occurs in all the N Atlantic arctic areas. (Elven)


The Epilobium hornemannii aggregate (E. behringianum, E. hornemannii, E. sertulatum)

Comments:

(1) Skvortsov synonymises in his draft, e.g., E. behringianum with E. hornemannii. My opinion, based only on the descriptions of E. behringianum and E. sertulatum, is that they must be more thoroughly evaluated before they can be reduced to synonymy. They are both kept as separate species by Hultén (1968) and Löve & Löve (1975). They are therefore tentatively treated as separate entities also here. (Elven)
70.1.5  Epilobium hornemannii Rchb. (1824), Pl. Crit. 73, t. 180.

S

2n= 36 (6x?).



2nD Löve & Löve (1975) list several counts, three as arctic.

G ICE NOR RUS RFE ALA CAN GRL

Comments:
70.1.6  Epilobium behringianum Hausskn. (1884), Monogr. Epilob. 277.

S

2n=



2nD

G ALA


Comments:

(1) Epilobium behringianum only reaches the Arctic in St. Matthew Isl., acc. to Hultén (1968). (Elven)


70.1.7  Epilobium sertulatum Hausskn. (1879), Oesterr. Bot. Zeitschr. 29: 52.

S

T [Described from Kamtchatka: Mutnaja Bay.]



2n=

2nD


G ALA

Comments:


70.1.8  Epilobium lactiflorum Hausskn. (1879), Oesterr. Bot. Zeitschr. 29: 89.

S

2n= 36 (6x?).



2nD Löve & Löve (1975) list four counts, two as arctic.

G ICE NOR RUS CAN GRL

Comments:
70.1.9  Epilobium anagallidifolium Lam. (1786), Encycl. 2: 376.

S E. alpinum L. (1753), nom. ambig., Sp. Pl. 348.

T [Described from France: "Mont-d'Or".]

2n= 36 (6x?).

2nD Löve & Löve (1975) list numerous counts, several as arctic.

G ICE NOR RUS SIB RFE ALA CAN GRL

Comments:

(1) Skvortsov (in draft and elsewhere) accepts E. alpinum as valid name of this taxon. I have here followed the current European and American usage and rejected the name E. alpinum. The diagnosis is insufficient for identification with any exact species, Linnaeus included three species in his 'alpinum' concept (anagallidifolium, alsinifolium and hornemannii), and an unambiguous typification seems to be lacking. Flora Europaea 2 (1968) cites, as synonym of E. anagallidifolium, "E. alpinum auct. mult. non L.". An unambiguous lectotypification and very good arguments are needed for re-introduction of that name as the valid one for this species. (Elven)


The Epilobium adenocaulon aggregate (E. adenocaulon/watsonii, E. ciliatum)

Comments:

(1) Two entities of this complex seem to reach the Arctic in America, and three entities (also E. glandulosum sensu scand.) are expanding northwards with an extraordinary speed as ruderals in W and N Fennoscandia. They may reach the Arctic as stable before the checklist is finished. I know that Skvortsov has made a study of these and there must be some correspondence between the names we apply in NW Europe (the ones above) and those applied in NE Europe (E. adenocaulon Hausskn. 1879, E. pseudorubescens Skvorts. 1995, and E. bergianum Skvorts. 1995). The 'watsonii' name is, e.g., applied now in Norway and Iceland but is not among those considered by Skvortsov in Tzvelev (1996, Fl. Vost. Evr. 9). It seems to have priority before E. adenocaulon and is also accepted by Scoggan (1979). There is very much confusion concerning the name(s) to apply. As these species are North American we should look for accepted American usage. (Elven)
70.1.10  Epilobium adenocaulon Hausskn. (1879), Oesterr. Bot. Zeitschr. 29: 119.

S E. glandulosum Lehm. [in Hook. (1832), Fl. Bor. Amer. 1: 206] var. adenocaulon (Haussk.) Fern. (1918), Rhodora 20: 35; E. watsonii Barbey in Brewer & Wats. (1876), Bot. Calif. 1: 219; E. ciliatum auct, non Raf. (1808).

2n= 36 (6x?).

2nD Löve & Löve (1975) list five non-arctic counts.

G ALA CAN

Comments:

(1) Added to Skvortsov's draft as it reaches the Arctic in several places, probably mostly as a ruderal but seed-reproducing and stable. The name E. watsonii Barbey is now usually used for this species in NW Europe. (Elven)
70.1.11  Epilobium ciliatum Raf. (1808), Med. Repos. 2, 5: 361.

S E. saximontanum Hausskn. (1879), Oesterr. Bot. Zeitschr. 29: 119; E. americanum Hausskn. (1879), Oesterr. Bot. Zeitschr. 29: 118; E. rubescens Rydb. (1904), Bull. Torrey Bot. Club 1904: 568.

2n=

2nD


G CAN

Comments:

(1) Added to Skvortsov's draft as it seems to reach the Arctic at least in the Hudson Bay area and as it at least has the potential to be stable. It is abundantly seed-reproducing and spreading just now in N Fennoscandia (like E. adenocaulon/watsonii and E. glandulosum sensu scand.).
70.1.12  Epilobium collinum C.C. Gmel. (1826), Fl. Bad. 4: 265.

S

2n= 36 (6x?).



2nD Löve & Löve (1975) list seven counts, one Icelandic.

G ICE


Comments:

(1) Added to Skvortsov's draft as it reaches the arctic parts of Iceland, acc. to Kristinsson's list and maps. (Elven)



70.2  Chamerion (Raf.) Raf. (1833), Herb. Raf. 51.

B Epilobium L. subgen. Chamerion Raf. (1818), Amer. Monthly Mag. & Crit. Rev. 2: 266.

S Chamaenerion Séguier (1754), nom. illegit., Pl. Veron. 3: 168.
70.2.1  Chamerion angustifolium (L.) Holub (1972), Folia Geobot. Phytotax. 7: 86.

B Epilobium angustifolium L. (1753), Sp. Pl. 347.

S Chamaenerion angustifolium (L.) Scop. (1772), nom. illegit., Fl. Carniol., ed. 2, 1: 271; Epilobium angustifolium L. subsp. angustifolium; Epilobium angustifolium L. subsp. macrophyllum (Hausskn.) Hultén (1967), Ark. Bot., ser. 2, 7: 85; Chamaenerion angustifolium (L.) Scop. subsp. macrophyllum (Hausskn.) S.K. Czerep. (1995), Vasc. Pl. Russia & Adj. States (former USSR) 319. (RFE? ALA); Chamerion platyphyllum (Daniels) Á. Löve & D. Löve (1976), Bot. Not. 128: 516 [basionym: Chamaenerion angustifolium (L.) Scop. var. platyphyllum Daniels (1911), Univ. Miss. Stud. 2, 2: 176].

2n= (1) 36 (6x?). (2) 72 (12x?).

2nD (1) Löve & Löve (1975) list numerous counts, a few as arctic, mainly in their subsp. intermedium (Wormskj.) Á. Löve ***; Lövkvist & Hultgård (1999 S Sweden). (2) Löve & Löve (1975) list eight non-arctic counts (for their C. platyphyllum).

G ICE NOR RUS SIB RFE ALA CAN GRL

Comments:

(1) Comments needed on subspecies and on C. platyphyllum (Daniels) Á. Löve & D. Löve with 2n=72. Is the Löve taxon the same as Hultén's subsp. macrophyllum? From Hultén's map it seems so. Löve & Löve (1976) describe it as: "... the octoploid (2n=72) taxon of southern boreal mountains in eastern North America and the western mountains north to the arctic region ...". The western North American plants look rather different from, e.g., the European ones but I am not sure they reach the Arctic as we define it. (Elven)



WARNING! Might be split on two entities with different ploidy levels.
70.2.2  Chamerion latifolium (L.) Holub (1972), Folia Geobot. Phytotax. 7: 86.

B Epilobium latifolium L. (1753), Sp. Pl. 347.

S Chamaenerion latifolium (L.) Th. Fr. & Lange (1877), Fl. Dan. 49: t. 2901, 2902; C. subdentatum (Rydb.) Löve & Löve (1976), Bot. Not. 128: 516 [basionym: Chamaenerion subdentatum Rydb. (1917), Fl. Rocky Mts. 585].

2n= (1) 36 (6x?). (2) 72 (12x?).

2nD (1) Löve & Löve (1975) list five counts, four as arctic (for their C. subdentatum). (2) Löve & Löve (1975) list several counts, nearly all as arctic.

G ICE RUS SIB RFE ALA CAN GRL

Comments:

(1) Comments needed on C. subdentatum (Rydb.) Á. Löve & D. Löve. Their comment is: "This is the tetraploid mainly western American and eastern Asiatic alpine taxon corresponding to the more widespread and almost circumpolar arctic-alpine octoploid species C. latifolium (L.) Holub.". (Elven)



WARNING! Might be split on two entities with different ploidy levels.

70.3  Circaea L. (1753), Sp. Pl. 9.

Comments:

(1) Genus and species tentatively added to Skvortsov's draft. (Elven)
70.3.1  Circaea alpina L. (1753), Sp. Pl. 9.

S

2n=



2nD

G ALA? CAN?

Comments:

(1) Approaches or reaches the Arctic on the east side of Hudson Bay ('Baffin-Labrador'), see Hultén & Fries (1986 map 1353), and also in the Yukon River delta, see Hultén (1968). A border case. (Elven)



71  Haloragaceae (Draft: R. Elven)

THIS DRAFT HAS NOT BEEN DISCUSSED

71.1  Myriophyllum L. (1753), Sp. Pl. 992.
71.1.1  Myriophyllum alterniflorum DC. in Lam. & DC. (1815), Fl. France, ed. 3, 5: 529.

S

T [Described from France: "dans la rivière d'Erdre pres Nantes".]



2n= 14 (2x).

2nD Löve & Löve (1975) list several counts, three as arctic.

G ICE NOR RUS ALA? CAN GRL

Comments:

(1) Not given for Alaska by Hultén (1968) but mapped from the Kotzebue area by Porsild & Cody (1980), probably just outside (east of) our arctic boundary. (Elven)
71.1.2  Myriophyllum verticillatum L. (1753), Sp. Pl. 992.

S

2n= 28 (4x).



2nD Löve & Löve (1975) list four non-arctic counts.

G RUS SIB? CAN

Comments:
71.1.3  Myriophyllum sibiricum Kom. (1914), Feddes Repert. 13: 168.

S M. exalbescens Fern. (1919), Rhodora 21: 120; M. spicatum L. subsp. exalbescens (Fern.) E. Murray (1982), Kalmia 12: 22; M. spicatum auct., non L. (1753).

2n= 42 (6x).

2nD Löve & Löve (1975) list three non-arctic counts.

G ICE? NOR? RUS SIB RFE ALA CAN GRL

Comments:



(1) Myriophyllum spicatum is given in Kristinsson's list as a border case in Iceland. It is here assumed that the plants belong to M. sibiricum but Icelandic occurrence of M. spicatum L. (1753), Sp. Pl. 992, s. str., cannot be discounted without a critical control of the material. (Elven)


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