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Comments: (1) Two subgeneric levels are applied in Yurtsev's draft: subgenus and section. Only two subgenera are given subg


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60.6  Oxytropis DC. (1802), Astragalogia 66 [or: p. 53?].

Comments:

(1) Two subgeneric levels are applied in Yurtsev's draft: subgenus and section. Only two subgenera are given   subg. Phacoxytropis Bunge (1874), Mém. Acad. Sci. Pétersb. (Sci. Phys. Math.), sér. 7, 22, 1: 6, with sect. Mesogaea and subg. Oxytropis (subg. Euoxytropis (Boiss.) Bunge (1874), Mém. Acad. Sci. Pétersb. (Sci. Phys. Math.), sér. 7, 22, 1: 47) with the others. As the subgenera do not differentiate, they are omitted. (Elven)

(2) The current Russian and North American treatments, exemplified by Yurtsev's draft and Gillett et al. (1999 onwards), diverge (if possible) even more in this genus than in Astragalus. There is much variation both in N Asia and N America but it is now treated quite differently. We have to reach some middle way between the extensive splitting in Russia-Sibera and the equally extensive lumping in N America. A problem is that the documentation of the Russian taxa is nearly inaccessible to us who do not read Russian. We also have very few specimens available for examination and evaluation. The treatment is therefore here preliminarily left more or less as it came from Yurtsev. However, all taxa not convincingly justified will automatically disappear into synonymy in the next version. (Elven)


Sect. Mesogaea Bunge (1874), Mém. Acad. Sci. Pétersb. (Sci. Phys. Math.), sér. 7, 22, 1: 38.
60.6.1  Oxytropis deflexa (Pall.) DC. (1802), Astragalogia 77.

B Astragalus deflexus Pall. (1779), Acta Horti Petrop. 2: 268.

S

2n= 16 (2x).



2nD Knaben & Engelskjøn (1967 N Norw, subsp. norvegica).

T Siberia: "ad nivalia juga Dauria ... in excelsis montibus circa Balyra rivum, aliosque Ononem influentibus", leg. S.P. Pallas (BM) holotype.

Comments:

(1) Flora Europaea gives for O. deflexa Astragalogia p. 96, Gillet et al. (1999 onward) p. 33. Hultén (1947: 1094) gives the year 1783 for Pallas' publication of Astragalus deflexus. (Elven)

(2) Several N American authors now refuse to accept subspecies (or other subspecific taxa), e.g., Cronquist et al. (1977, Interm. Fl.), Isely (1984, Iowa St. J. Res. 59) and Gillett et al. (1999 onwards). I am, however, aware of fairly distinct differences between N European, Asian and N American plants and can accept some races. There is also chemical support of some differentiation (**REF**).

Hultén & Fries (1986) divided the variation into three subspecies: subsp. deflexa in C and NE Asia east to Kolyma, subsp. retrorsa (Fern.) Hultén ined. (same as subsp. foliolosa) in western N America, and subsp. norvegica Nordh. in a very small and isolated (non-arctic) area in N Norway. The fourth subspecies below, subsp. dezhnevii, is obviously based on new finds in Russian Far East, in the gap between subsp. deflexa and subsp. foliolosa. It needs special justification as a separate taxon and not only as a transition. (Elven)



WARNING! Subspecies might be omitted.
60.6.1.1  Oxytropis deflexa (Pall.) DC. subsp. deflexa

S

2n= 16 (2x).



2nD Yurtsev's draft.

G SIB


Comments:
60.6.1.2  Oxytropis deflexa (Pall.) DC. subsp. dezhnevii (Jurtz.) Jurtz. (1986), Fl. Arct. URSS 9, 2: 78.

B O. deflexa (Pall.) DC. var. dezhnevii Jurtz. (1972), Bot. Zhurn. 57,6: 647.

S

2n= 16 (2x).



2nD Yurtsev & Tzvelev (1972 NE As).

G RFE


Comments:

(1) Fl. Arct. URSS maps it only from three scattered sites in Chukchi Peninsula. This is too little on which to base a regional race, i.e., subspecies. The distance from the nearest site in Alaska in westernmost Brooks Range is not insurmountable. (Elven)

(2) The count from NE Asia listed by Löve & Löve (1975) under subsp. foliolosa has been transferred here as this is the only race of O. deflexa reported from the area. (Elven)
60.6.1.3  Oxytropis deflexa (Pall.) DC. subsp. foliolosa (Hook.) Cody ***

B O. foliolosa Hook. (1834), Fl. Bor.-Amer. 1: 146.

S O. deflexa (Pall.) DC. var. foliolosa (Hook.) Barneby (1951), Leafl. West. Bot. 5: 111; O. deflexa (Pall.) DC. var. sericea Torr. & A. Gray ***

T Canada: Alberta, "Ox. foliolosa Hook. Fl. Bor. Amer. Rocky Mountains", leg. Drummond (K) holotype.

2n= 16 (2x).

2nD Löve & Löve (1975) list four non-arctic counts (see also subsp. dezhnevii).

G ALA CAN

Comments:

(1) Hultén's maps show a strange discrepancy. Hultén (1968) maps two N American races of O. deflexa, the western var. sericea Torr. & A. Gray that does not reach the Arctic and the disjunctly transcontinental var. foliolosa that does. Hultén & Fries (1986) exclude the eastern part areas from the species, without comment. Gillett et al. (1999 onwards) also map the eastern parts as it here reaches both northernmost Labrador and Ungava and S Baffin Island. Var. sericea has to be included in the synonymy as it is an accepted name in 1968. (Elven)
Sect. Arctobia Bunge (1874), Mém. Acad. Sci. Pétersb. (Sci. Phys. Math.), sér. 7, 22, 1: 112.

S Sect. Caeciabia Bunge (1874), Mém. Acad. Sci. Pétersb. (Sci. Phys. Math.), sér. 7, 22, 1: 115.


The Oxytropis nigrescens aggregate (O. arctobia, O. bryophila, O. czukotica, O. glaberrima, O. gorodkovii, O. kokrinensis, O. nigrescens, O. pumilio, ?O. sublongipes)

Comments:

(1) The North American view is to consider the entire American part of the aggregate as one variable species with 'varieties' (Barneby 1952, later taken up by Gillett et al. 1999 onwards). As the 'varieties' mostly occupy different regions, they would more naturally be subspecies in our concept. This is also supported by the ranges of the major entities: 'nigrescens' s. str. from Jenisei to Kolyma, 'czukotica' from Kolyma to the Bering Strait, 'pygmaea/gorodkovii' around the Bering Strait, 'bryophila' in Alaska-Yukon, and 'arctobia' in N Canada. They are all di  or tetraploid, parapatric and closely related. I am therefore much in favour of a 'reversal' to Hultén's opinion: subspecies for the major entities.

Two questions are: (a) How many subspecies; and (b) What to do about the 'minor' entities? There are three major entities on the North American side; 'arctobia' is comparatively distinct from 'bryophila' and also allopatric from it, and even if 'bryophila' and 'pygmaea' are mostly overlapping geographically, in my opinion they differ enough as to merit rank. Oxytropis glaberrima occurs (acc. to Hultén 1968) in a single site in Brooks Range and is said to differ from 'bryophila' only in being glabrous. It is obviously not a major variation and should perhaps not even be regarded as a variety but could well be result of a single mutation. It should be reduced to synonymy. Oxytropis kokrinensis is more problematic. In 1968 it was known from three sites only, within the range of 'bryophila' from which it differs in at least two assumed independent characters. It should be compared with the Asian entities.

On the Asian side, the map presented by Yurtsev (1986, Fl. Arct. URSS 9, 2) of O. nigrescens and O. czukotica indicate some close connection; the former entity is frequent straight up to a line where it 'changes' into the latter and equally frequent entity. This line runs from Stanovoy Khrebet Mts. near the Chinese border to just west of the Kolyma estuary, with not a single overlap. To a non-Russian this looks like a character transition in the west-east direction within a single species. Two subspecies may be acceptable, but not two species. Oxytropis pumilio is not treated by Fl. Arct. URSS but is a southern entity (subspecies or species) just penetrating our area in Russian Far East. It may be an Asian parallel to the more differentiated taxa (probably 'good' species) of the complex in the Rockies and Central Alaska: O. huddelsonii A.E.Porsild and O. podocarpa A. Gray. Oxytropis longipes in Russian Far East may be of the same type.

WARNING! Might be considered as one wide species with several subspecies and varieties.
60.6.2  Oxytropis nigrescens (Pall.) Fisch. in DC (1825), Prodr. 2: 278.

B Astragalus nigrescens Pall. (1800), Spec. Astrag. 65.

S O. nigrescens (Pall.) Fisch. subsp. nigrescens.

T [Described from: "inter Aldanum fl. et orientalem Oceanum", leg. D.D. Merk]

2n= 16 (2x).

2nD Holmen (1962), but see below; Ledingham & Reever (1963); Yurtsev & Zhukova (1968).

G SIB

Comments:



(1) Löve & Löve (1975) list the three abovementioned counts for O. nigrescens s. str., but as two of them are North American, the 'sensu stricto' of the Löves is not nearly as strict as Yurtsev's. See also Yurtsev's comment on the Holmen count under O. kokrinensis. (Elven)

WARNING! This entity should probably be considered the type subspecies of a wider species.
60.6.3  Oxytropis czukotica Jurtz. (1980), Byull. Mosk. Obshch. Ispyt. Prir., Otd. Biol. 85, 6: 102.

S O. tschuktschorum Jurtz. (1968), Bot. Zhurn. 53: 1538, p. max. p., exclus. typo.

2n= 32 (4x).

2nD Yurtsev's draft.

G SIB RFE ALA

Comments:

(1) The only locality in North America is Ogotoruk Creek at Camp Thompson in northwesternmost Alaska. (Yurtsev)

(2) As NW Alaska is the area where also the 'bryophila', 'pygmaea' and 'kokrinensis' entities are known, the Ogotoruk Creek material should be very critically studied before a fourth entity is accepted from there. (Elven)



WARNING! Should probably either be reduced to a subspecies of O. nigrescens or merged fully with O. nigrescens subsp. nigrescens as part of an east-west cline.
60.6.4  Oxytropis pumilio (Pall.) Ledeb. (1842), Fl. Ross. 1: 589.

B Astragalus pumilio Pall. (1800), Spec. Astrag. 67, t. 55.

S

T [Described from the Kuriles]



2n= 16 (2x).

2nD Yurtsev's draft.

G RFE

Comments:



(1) Chromosome counts from the northernmost Kuriles and southernmost Kamtchatka. (Yurtsev)

(2) The only specimen from the Panarctic Flora area was collected on the northern Koryak Coast near Meinopylgeno Village (specimen incomplete). (Yurtsev)

(3) May merit status as a separate species. (Elven)
60.6.5  Oxytropis bryophila (Greene) Jurtz. (1968), Bot. Zhurn. 53: 1539.

B Aragallus bryophilus Greene (1905), Proc. Biol. Soc. Wash. 18: 17.

S O. nigrescens (Pall.) Fisch. subsp. bryophila (Greene) Hultén (1947), Lunds Univ. Årsskr., n. f., avd. 2, 43, 1: 1102.

T Alaska: Hall I., 11.08.1891, leg. Macoun (CAN) holotype?, see comment.

2n= (1) 16 (2x). (2) 32? (4x).

2nD ?


G RFE ALA CAN

Comments:

(1) Hultén (1947) states that the entity is described from St. Matthew Island but cites a type from Hall Island. (Elven)

(2) Yurtsev reports this entity also from Russian Far East in his draft. This means that three entities co-occur in Chukchi Peninsula: 'czukotica', 'bryophila' and 'gorodkovii/pygmaea'. It may be a result of duplicate naming on the Russian and American side and therefore one too many. (Elven)



WARNING! Should probably be reduced to a subspecies of O. nigrescens.
60.6.6  Oxytropis glaberrima Hultén (1954), Bot. Tidsskr. 51: 124.

S

2n=



2nD

G ALA


Comments:

(1) The status of the taxon needs a further examination. It my be a glabrous race (form?) of O. bryophila. (Yurtsev)

(2) See comments to the aggregate. I agree fully with Yurtsev; form may be the best category. It is certainly not a 'race'. (Elven)

WARNING! Should be reduced to synonymy within O. nigrescens subsp. bryophila.
60.6.7  Oxytropis kokrinensis A.E. Porsild (1939), Rhodora 41: 251.

S

T Alaska: Kokrines Mts., leg. Porsild 762 (CAN) holotype.



2n= 16 (2x).

2nD Yurtsev's draft.

G ALA

Comments: See comments under the aggregate.



(1) The voucher specimen for K.Holmen's counting 2n=16 for O. nigrescens (Holmen in Bot. Not. 115: 90) contains both O. bryophila and O. kokrinensis, but no O. nigrescens. (Yurtsev)

WARNING! Might be reduced to minor variation in one of the proposed subspecies.
60.6.8  Oxytropis gorodkovii Jurtz. (1968), Bot. Zhurn. 53: 1538.

S Astragalus pygmaeus Pall. (1800), Spec. Astragal. 66; O. nigrescens (Pall.) Fisch. subsp. pygmaea (Pall.) Hultén (1947), Lunds Univ. Årsskr., n. f., avd. 2, 43, 1: 1103; O. pygmaea (Pall.) Fern. (1928), Rhodora 30: 153, non Tausch ex G. Beck (1901); O. tschukschorum Jurtz. (1968), Bot. Zhurn. 53: 1538, p. min. p., quoad typo.

2n= (1) 16 (2x). (2) 32 (4x).

2nD (1) Johnson & Packer (1968 Ala); Yurtsev & Zhukova (1968 NE As); Zhukova & Petrovsky (1971 NE As).

G RFE ALA

Comments:



WARNING! Should probably be reduced to a subspecies as O. nigrescens subsp. pygmaea.
60.6.9  Oxytropis arctobia Bunge (1871), Mem. Acad. Petersb., ser. 7, 22, 1: 114.

S O. nigrescens (Pallas) Fisch. ex DC. subsp. arctobia (Bunge) Hultén (1968), Ark. Bot., ser. 2, 7, 1: 80; O. nigrescens (Pallas) Fisch. ex DC. var. uniflora (Hook.) Barneby (1952), Proc. Calif. Acad. Sci., ser. 4, 27: 209.

2n= 16 (2x).

2nD Gillett et al. (1999 secondary ref.).

G CAN

Comments:



(1) Note that Gillett et al. (1999), in the most recent Canadian treatise of arctic legumes, treat this as a variety of O. nigrescens. (Elven)

WARNING! Should probably be reduced to a subspecies as O. nigrescens subsp. arctobia.
60.6.10  Oxytropis sublongipes Jurtz. (1979), Byull. Mosk. Obshch. Ispyt. Prir., Otd. Biol. 84, 1: 99.

S O. kamtschatica auct., non Hultén (1929).

2n= 16 (2x).

2nD Yurtsev's draft.

G RFE

Comments:



(1) I need to know more about this entity before a proposal can be made. No material is available here in Oslo. (Elven)

WARNING! Should be re-evaluated as a possible subspecies of O. nigrescens s. lat. or as part of a O. revoluta group.
60.6.11  Oxytropis revoluta Ledeb. (1842), Fl. Ross. 1: 588.

S

2n= 16 (2x).



2nD Sokolovskaya (1963, 1965).

G RFE


Comments:

(1) Seems to belong to a 'Ochotian' species aggregate that also includes O. exserta Jurtz. 1979 and O. sublongipes Jurtz. 1979. As O. revoluta is the oldest name in this group it is probably suitable also for a more widely considered species. (Elven)


60.6.12  Oxytropis podocarpa A. Gray (1864), Proc. Amer. Acad. 6: 234.

S

T Canada: Labrador, leg. Schweinitz (???) lectotype, selected by Fernald (1928).



2n= 16 (2x).

2nD Ledingham (1960).

G CAN

Comments:



(1) Chromosome number unknown, acc. to Gillett et al. (1999). (Elven)
60.6.13  Oxytropis scammaniana Hultén (1946), Ark. Bot. 33 B, 1: 4.

S

T Alaska: Eagle Summit, leg. Scamman 806 (G, H).



2n=

2nD


G ALA

Comments:


60.6.14  Oxytropis mertensiana Turcz. (1840), Bull. Soc. Naturalistes Moscou 13: 68.

S

T [Described from: Chukchi peninsula, St. Lawrence Bay]



2n= 16 (2x).

2nD Löve & Löve (1975) list eight American and Russian counts, all as arctic.

G SIB RFE ALA

Comments:



(1) Hultén (1947: 1101) gives page 58 for the publication of the name. Russians, please check. (Elven)


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